This is a companion discussion topic for the original entry at https://www.kirupa.com/animations/customizing_css_animations.htm
A TL;DR:
I was hoping that this trick would allow me to have a random circle moving around the screen, fully powered by a CSS animation only. As it turns out, that is possible…albeit with some major quirks.
Here is an example:
This only works in Chrome stable (as of now). It doesn’t work in Chrome Canary, Firefox, or Safari. In the keyframes, do not remove the background-color declaration. For some reason, it needs to be there to get the circle to move. No idea why 
And…the video for it is up as well:
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